Stomiidae

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Stomiidae is a family of deep-sea ray-finned fish including the barbeled dragonfishes and (in the subfamily Malacosteinae) the loosejaws.

Species

There are 276 species in 27 genera:

• Genus Aristostomias
• * Aristostomias grimaldii Zugmayer, 1913.
• * Aristostomias lunifer Regan & Trewavas, 1930.
• * Aristostomias polydactylus Regan & Trewavas, 1930.
• * Shiny loosejaw, Aristostomias scintillans (Gilbert, 1915).
• * Aristostomias tittmanni Welsh, 1923.
• * Aristostomias xenostoma Regan & Trewavas, 1930.
• Genus Astronesthes
• * Astronesthes atlanticus Parin & Borodulina, 1996.
• * Astronesthes bilobatus Parin & Borodulina, 1996.
• * Astronesthes boulengeri Gilchrist, 1902.
• * Astronesthes caulophorus Regan & Trewavas, 1929.
• * Astronesthes chrysophekadion (Bleeker, 1849).
• * Astronesthes cyaneus (Brauer, 1902).
• * Astronesthes cyclophotus Regan & Trewavas, 1929.
• * Astronesthes decoratus Parin & Borodulina, 2002.
• * Astronesthes dupliglandis Parin & Borodulina, 1997.
• * Astronesthes exsul Parin & Borodulina, 2002.
• * Astronesthes fedorovi Parin & Borodulina, 1994.
• * Astronesthes galapagensis Parin, Borodulina & Hulley, 1999.
• * Snaggletooth, Astronesthes gemmifer Goode & Bean, 1896.

Snaggletooth, Astronesthes gemmifer

• * Astronesthes gibbsi Borodulina, 1992.
• * Astronesthes gudrunae Parin & Borodulina, 2002.
• * Astronesthes haplophos Parin & Borodulina, 2002.
• * Astronesthes ijimai Tanaka, 1908.
• * Astronesthes illuminatus Parin, Borodulina & Hulley, 1999.
• * Astronesthes indicus Brauer, 1902.
• * Astronesthes indopacificus Parin & Borodulina, 1997.
• * Astronesthes karsteni Parin & Borodulina, 2002.
• * Astronesthes kreffti Gibbs & McKinney, 1988.
• * Astronesthes lamellosus Goodyear & Gibbs, 1970.
• * Astronesthes lampara Parin & Borodulina, 1998.
• * Astronesthes leucopogon Regan & Trewavas, 1929.
• * Astronesthes longiceps Regan & Trewavas, 1929.
• * Astronesthes lucibucca Parin & Borodulina, 1996.
• * Pacific astronesthid fish, Astronesthes lucifer Gilbert, 1905.
• * Astronesthes luetkeni Regan & Trewavas, 1929.
• * Astronesthes lupina Whitley, 1941.
• * Astronesthes macropogon Goodyear & Gibbs, 1970.
• * Astronesthes martensii Klunzinger, 1871.
• * Astronesthes micropogon Goodyear & Gibbs, 1970.
• * Astronesthes neopogon Regan & Trewavas, 1929.
• * Astronesthes niger Richardson, 1845.
• * Astronesthes nigroides Gibbs & Aron, 1960.
• * Astronesthes oligoa Parin & Borodulina, 2002.
• * Astronesthes psychrolutes (Gibbs & Weitzman, 1965).
• * Astronesthes quasiindicus Parin & Borodulina, 1996.
• * Astronesthes richardsoni (Poey, 1852).

Astronesthes richardsoni

• * Astronesthes similus Parr, 1927.
• * Astronesthes spatulifer Gibbs & McKinney, 1988.
• * Astronesthes splendida Brauer, 1902.
• * Astronesthes tanibe Parin & Borodulina, 2001.
• * Astronesthes tatyanae Parin & Borodulina, 1998.
• * Astronesthes tchuvasovi Parin & Borodulina, 1996.
• * Astronesthes trifibulatus Gibbs, Amaoka & Haruta, 1984.
• * Astronesthes zetgibbsi Parin & Borodulina, 1997.
• * Astronesthes zharovi Parin & Borodulina, 1998.
• Genus Bathophilus
• * Bathophilus abarbatus Barnett & Gibbs, 1968.
• * Bathophilus altipinnis Beebe, 1933.
• * Bathophilus ater (Brauer, 1902).
• * Bathophilus brevis Regan & Trewavas, 1930.
• * Bathophilus digitatus (Welsh, 1923).
• * Sparing, Bathophilus filifer (Regan & Trewavas, 1929).
• * Highfin dragonfish, Bathophilus flemingi Aron & McCrery, 1958.
• * Bathophilus indicus (Brauer, 1902).
• * Bathophilus irregularis Norman, 1930.
• * Bathophilus kingi Barnett & Gibbs, 1968.
• * Bathophilus longipinnis (Pappenheim, 1914).
• * Scaleless dragonfish, Bathophilus nigerrimus Giglioli, 1882.

Scaleless dragonfish, Bathophilus nigerrimus

• * Bathophilus pawneei Parr, 1927.
• * Bathophilus proximus Regan & Trewavas, 1930.
• * Bathophilus schizochirus Regan & Trewavas, 1930.
• * Bathophilus vaillanti (Zugmayer, 1911).
• Genus Borostomias
• * Borostomias abyssorum (Koehler, 1896).
• * Borostomias antarcticus (Lönnberg, 1905).
• * Borostomias elucens (Brauer, 1906).
• * Borostomias mononema (Regan & Trewavas, 1929).
• * Borostomias pacificus (Imai, 1941).
• * Panama snaggletooth, Borostomias panamensis Regan & Trewavas, 1929.
• Genus Chauliodus
• * Chauliodus barbatus (Lowe, 1843).
• * Dana viperfish, Chauliodus danae Regan & Trewavas, 1929.

Dana viperfish, Chauliodus danae

• * Chauliodus dentatus Garman, 1899.
• * Pacific viperfish, Chauliodus macouni Bean, 1890.
• * Chauliodus minimus Parin & Novikova, 1974.
• * Chauliodus pammelas Alcock, 1892.
• * Chauliodus schmidti (Regan & Trewavas, 1929).
• * Sloane's viperfish, Chauliodus sloani Bloch & Schneider, 1801.

Sloane's viperfish, Chauliodus sloani

• * Chauliodus vasnetzovi Novikova, 1972.
• Genus Chirostomias
• * Chirostomias pliopterus Regan & Trewavas, 1930.
• Genus Echiostoma
• * Echiostoma barbatum Lowe, 1843.

Echiostoma barbatum

• Genus Eupogonesthes
• * Eupogonesthes xenicus Parin & Borodulina, 1993.
• Genus Eustomias
• * Eustomias achirus Parin & Pokhil'skaya, 1974.
• * Eustomias acinosus Regan & Trewavas, 1930.
• * Eustomias aequatorialis Clarke, 1998.
• * Eustomias albibulbus Clarke, 2001.
• * Eustomias appositus Gibbs, Clarke & Gomon, 1983.
• * Eustomias arborifer Parr, 1927.
• * Eustomias australensis Gibbs, Clarke & Gomon, 1983.
• * Eustomias austratlanticus Gibbs, Clarke & Gomon, 1983.
• * Eustomias bertelseni Gibbs, Clarke & Gomon, 1983.
• * Eustomias bibulboides Gibbs, Clarke & Gomon, 1983.
• * Eustomias bibulbosus Parr, 1927.
• * Eustomias bifilis (Regan & Trewavas, 1929).
• * Eustomias bigelowi Welsh, 1923.
• * Eustomias bimargaritatus Regan & Trewavas, 1930.
• * Eustomias bimargaritoides Gibbs, Clarke & Gomon, 1983.
• * Eustomias binghami Parr, 1927.
• * Eustomias bituberatus Regan & Trewavas, 1930.
• * Eustomias bituberoides Gibbs, Clarke & Gomon, 1983.
• * Eustomias borealis Clarke, 2000.
• * Eustomias braueri (Regan, 1908).
• * Eustomias brevibarbatus Parr, 1927.
• * Eustomias bulbiramis Clarke, 2001.
• * Eustomias bulbornatus Gibbs, 1960.
• * Eustomias cancriensis Gibbs, Clarke & Gomon, 1983.
• * Eustomias cirritus Gibbs, Clarke & Gomon, 1983.
• * Eustomias contiguus Gomon & Gibbs, 1985.
• * Eustomias crossotus Gibbs, Clarke & Gomon, 1983.
• * Eustomias crucis Gibbs & Craddock, 1973.
• * Eustomias cryptobulbus Clarke, 2001.
• * Eustomias curtatus Gibbs, Clarke & Gomon, 1983.
• * Eustomias curtifilis Clarke, 2000.
• * Eustomias danae (Regan & Trewavas, 1929).
• * Eustomias decoratus (Parin & Borodulina, 2002).
• * Eustomias dendriticus Regan & Trewavas, 1930.
• * Eustomias deofamiliaris Gibbs, Clarke & Gomon, 1983.
• * Eustomias digitatus (Welsh, 1923).
• * Eustomias dinema Clarke, 1999.
• * Eustomias dispar Gomon & Gibbs, 1985.
• * Eustomias dubius Parr, 1927.
• * Eustomias elongatus Clarke, 2001.
• * Eustomias enbarbatus Welsh, 1923.
• * Eustomias filifer (Regan & Trewavas, 1929).
• * Eustomias fissibarbis (Pappenheim, 1914).
• * Eustomias flagellifer Clarke, 2001.
• * Eustomias furcifer Regan & Trewavas, 1930.
• * Eustomias gibbsi (Borodulina, 1992).
• * Eustomias grandibulbus Gibbs, Clarke & Gomon, 1983.
• * Eustomias hulleyi Gomon & Gibbs, 1985.
• * Eustomias hypopsilus Gomon & Gibbs, 1985.
• * Eustomias ignotus Gomon & Gibbs, 1985.
• * Eustomias inconstans Gibbs, Clarke & Gomon, 1983.
• * Eustomias insularum Clarke, 1998.
• * Eustomias intermedius Clarke, 1998.
• * Eustomias interruptus Clarke, 1999.
• * Eustomias ioani Parin & Pokhil'skaya, 1974.
• * Eustomias jimcraddocki Sutton & Hartel, 2004.
• * Eustomias kreffti (Gibbs & McKinney, 1988).
• * Eustomias lanceolatus Clarke, 1999.
• * Eustomias leptobolus Regan & Trewavas, 1930.
• * Eustomias lipochirus Regan & Trewavas, 1930.
• * Eustomias longibarba Parr, 1927.
• * Eustomias longiramis Clarke, 2001.
• * Eustomias macronema Regan & Trewavas, 1930.
• * Eustomias macrophthalmus (Regan & Trewavas, 1929).
• * Eustomias macrurus Regan & Trewavas, 1930.
• * Eustomias magnificus Clarke, 2001.
• * Eustomias medusa Gibbs, Clarke & Gomon, 1983.
• * Eustomias melanonema Regan & Trewavas, 1930.
• * Eustomias melanostigma Regan & Trewavas, 1930.
• * Eustomias melanostigmoides Gibbs, Clarke & Gomon, 1983.
• * Eustomias mesostenus Gibbs, Clarke & Gomon, 1983.
• * Eustomias metamelas Gomon & Gibbs, 1985.
• * Eustomias micraster Parr, 1927.
• * Eustomias micropterygius Parr, 1927.
• * Eustomias minimus (Parin & Novikova, 1974).
• * Eustomias monoclonoides Clarke, 1999.
• * Eustomias monoclonus Regan & Trewavas, 1930.
• * Eustomias monodactylus Regan & Trewavas, 1930.
• * Eustomias multifilis Parin & Pokhil'skaya, 1978.
• * Eustomias obscurus Vaillant, 1884.

Eustomias obscurus

• * Eustomias orientalis Gibbs, Clarke & Gomon, 1983.
• * Eustomias pacificus (Imai, 1941).
• * Eustomias parini Clarke, 2001.
• * Eustomias parri Regan & Trewavas, 1930.
• * Eustomias patulus Regan & Trewavas, 1930.
• * Eustomias paucifilis Parr, 1927.
• * Eustomias paxtoni Clarke, 2001.
• * Eustomias perplexus Gibbs, Clarke & Gomon, 1983.
• * Eustomias pinnatus Clarke, 1999.
• * Eustomias polyaster Parr, 1927.
• * Eustomias posti Gibbs, Clarke & Gomon, 1983.
• * Eustomias precarius Gomon & Gibbs, 1985.
• * Eustomias problematicus Clarke, 2001.
• * Eustomias pyrifer Regan & Trewavas, 1930.
• * Eustomias quadrifilis Gomon & Gibbs, 1985.
• * Eustomias radicifilis Borodin, 1930.
• * Eustomias satterleei Beebe, 1933.
• * Eustomias schiffi Beebe, 1932.
• * Eustomias schmidti (Regan & Trewavas, 1929).
• * Eustomias silvescens Regan & Trewavas, 1930.
• * Eustomias similis (Parr, 1927).
• * Eustomias simplex (Parr, 1927).
• * Eustomias spherulifer Gibbs, Clarke & Gomon, 1983.
• * Eustomias suluensis Gibbs, Clarke & Gomon, 1983.
• * Eustomias tenisoni Regan & Trewavas, 1930.
• * Eustomias tetranema Zugmayer, 1913.
• * Eustomias teuthidopsis Gibbs, Clarke & Gomon, 1983.
• * Eustomias tomentosis Clarke, 1998.
• * Eustomias trewavasae Norman, 1930.
• * Eustomias triramis Regan & Trewavas, 1930.
• * Eustomias uniramis Clarke, 1999.
• * Eustomias variabilis Regan & Trewavas, 1930.
• * Eustomias vitiazi Parin & Pokhil'skaya, 1974.
• * Eustomias vulgaris Clarke, 2001.
• * Eustomias woollardi Clarke, 1998.
• * Eustomias xenobolus Regan & Trewavas, 1930.
• Genus Flagellostomias
• * Flagellostomias boureei (Zugmayer, 1913).
• Genus Grammatostomias
• * Grammatostomias circularis Morrow, 1959.
• * Grammatostomias dentatus (Garman, 1899).

Grammatostomias dentatus

• * Grammatostomias flagellibarba Holt & Byrne, 1910.
• Genus Heterophotus
• * Heterophotus ophistoma Regan & Trewavas, 1929.
• Genus Idiacanthus
• * Pacific blackdragon, Idiacanthus antrostomus Gilbert, 1890.
• * Idiacanthus atlanticus (Parin & Borodulina, 1996).
• * Ribbon sawtail fish, Idiacanthus fasciola Peters, 1877.

Ribbon sawtail fish, Idiacanthus fasciola

• Genus Leptostomias
• * Leptostomias analis Regan & Trewavas, 1930.
• * Leptostomias bermudensis Beebe, 1932.
• * Leptostomias bilobatus (Parin & Borodulina, 1996).
• * Leptostomias gladiator (Zugmayer, 1911).
• * Leptostomias gracilis Regan & Trewavas, 1930.
• * Leptostomias haplocaulus Regan & Trewavas, 1930.
• * Leptostomias leptobolus (Regan & Trewavas, 1930).
• * Leptostomias longibarba (Parr, 1927).
• * Long-threaded dragonfish, Leptostomias macronema (Regan & Trewavas, 1930).
• * Leptostomias macropogon (Goodyear & Gibbs, 1970).
• * Leptostomias multifilis (Parin & Pokhil'skaya, 1978).
• * Leptostomias robustus Imai, 1941.
• Genus Malacosteus
• * Malacosteus indicus (Brauer, 1902).
• * Stoplight loosejaw, Malacosteus niger (Ayres, 1848).

Stoplight loosejaw, Malacosteus niger

• Genus Melanostomias
• * Melanostomias bartonbeani Parr, 1927.
• * Melanostomias biseriatus Regan & Trewavas, 1930.
• * Melanostomias globulifer Fowler, 1934.
• * Melanostomias macrophotus Regan & Trewavas, 1930.
• * Melanostomias margaritifer Regan & Trewavas, 1930.
• * Melanostomias melanopogon Regan & Trewavas, 1930.
• * Melanostomias melanops Brauer, 1902.
• * Melanostomias niger (Richardson, 1845).
• * Melanostomias nigroaxialis Parin & Pokhil'skaya, 1978.
• * Melanostomias paucilaternatus Parin & Pokhil'skaya, 1978.
• * Melanostomias pauciradius Matsubara, 1938.
• * Melanostomias pollicifer Parin & Pokhil'skaya, 1978.
• * Melanostomias stewarti Fowler, 1934.
• * Melanostomias tentaculatus (Regan & Trewavas, 1930).
• * Valdivia black dragonfish, Melanostomias valdiviae Brauer, 1902.
• * Melanostomias vierecki Fowler, 1934.
• Genus Neonesthes
• * Neonesthes capensis (Gilchrist & von Bonde, 1924).
• * Neonesthes microcephalus Norman, 1930.
• Genus Odontostomias
• * Odontostomias masticopogon Norman, 1930.
• * Odontostomias micropogon (Goodyear & Gibbs, 1970).
• Genus Opostomias
• * Opostomias micripnus (Günther, 1878).

Opostomias micripnus

• * Pitgum lanternfish, Opostomias mitsuii Imai, 1941.
• Genus Pachystomias
• * Pachystomias microdon (Günther, 1878).

Pachystomias microdon

• Genus Photonectes
• * Photonectes achirus (Parin & Pokhil'skaya, 1974).
• * Photonectes albipennis (Döderlein, 1882).
• * Photonectes braueri (Regan, 1908).
• * Photonectes caerulescens Regan & Trewavas, 1930.
• * Photonectes dinema (Clarke, 1999).
• * Photonectes gracilis (Regan & Trewavas, 1930).

Photonectes gracilis

• * Photonectes leucospilus Regan & Trewavas, 1930.
• * Photonectes margarita (Goode & Bean, 1896).

Photonectes margarita

• * Photonectes mirabilis Parr, 1927.
• * Photonectes munificus Gibbs, 1968.
• * Photonectes parvimanus Regan & Trewavas, 1930.
• * Photonectes phyllopogon Regan & Trewavas, 1930.
• Genus Photostomias
• * Photostomias guernei Collett, 1889.

Photostomias guernei

• * Photostomias mirabilis (Parr, 1927).
• Genus Rhadinesthes
• * Rhadinesthes decimus (Zugmayer, 1911).
• Genus Stomias
• * Günther's boafish, Stomias affinis Günther, 1887.

Günther's boafish, Stomias affinis

• * Black-belly dragonfish, Stomias atriventer Garman, 1899.
• * Scaly dragonfish, Stomias boa boa (Risso, 1810).

Scaly dragonfish, Stomias boa boa

• * Stomias boa colubrinus Garman, 1899.
• * Boa dragonfish, Stomias boa ferox Reinhardt, 1842.

Boa dragonfish, Stomias boa ferox

• * Stomias brevibarbatus (Parr, 1927).
• * Stomias danae (Regan & Trewavas, 1929).
• * Stomias gracilis (Regan & Trewavas, 1930).
• * Stomias lampropeltis Gibbs, 1969.
• * Stomias longibarbatus (Brauer, 1902).
• * Alcock's boafish, Stomias nebulosus Alcock, 1889.
• Genus Tactostoma
• * Longfin dragonfish, Tactostoma macropus Bolin, 1939.
• Genus Thysanactis
• * Thysanactis dentex Regan & Trewavas, 1930.
• Genus Trigonolampa
• * Threelight dragonfish, Trigonolampa miriceps Regan & Trewavas, 1930.

References

• ["Stomiidae"]. FishBase. Ed. Ranier Froese and Daniel Pauly. January 2006 version. N.p.: FishBase, 2006.

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